Curriculum Vitae, Academic Record and Publication List of

Assoc. Prof. Levent Ozturk





Name                                                       : Levent Ozturk

Mailing Address                                   : Sabanci University

                    Faculty of Engineering and Natural Sciences

                                                    Istanbul-TURKEY (TR)

Place and Date of Birth                      : Mersin-Turkey, 16.03.1971

Residency Status                                   : Republic of Turkey

Gender                                                    : Male

Current Position                                   : Faculty Member

Martial Status                                       : Married

Phone                                                      : +90-216-483-9613

Fax                                                           : +90-216-483-9550

E-mail                                                     :





1990-1994     Bachelor’s degree from Cukurova University, Faculty of Agriculture, Department of Soil Science

1994-1996     MSc. degree from Cukurova University, Faculty of Agriculture, Department of Soil Science

10.03.1995     Employment as Research Assistant at Cukurova University, Institute of Natural and Applied Sciences

1996-2001     PhD. degree from Cukurova University, Institute of Natural and Applied Sciences

21.06.2001     Employment as an Assistant Professor at Cukurova University, Faculty of Agriculture

14.04.2004     Possession of Associate Professorship degree by ÜAK of Turkey

01.07.2004     Employment as a visiting scientist at Sabanci University, Faculty of Engineering and Natural Sciences

01.03.2006     Employment as a faculty member at Sabanci University, Faculty of Engineering and Natural Sciences (continuing)

27.09.2010-25.08.2011        Sabbatical leave for two semesters at the USDA Nutrition Lab, Cornell University USA





15.07-15.09.1994     Summer training at Hohenheim University, Institute for Plant Nutrition, Stuttgart-Germany

15.08-12.09.1996     Training on laboratory techniques at Hohenheim University, Institute for Plant Nutrition, Stuttgart-Germany

07.03-24.03.1997     Technical Course on ICP (Inductively Coupled Plasma Argon-Emission Spectrometer) operation at Jobin Yvon I.S.A., Paris-France

10.11-16.11.1997     Technical training on Photosynthesis Rate Analyzer at Analytical Development Co., London-UK

07.10-11.12.1998     Training on laboratory techniques at Weizmann Institute of Science, Plant Genetics Department, Rehovot-Israel

20.04-17.05.1999     Training on trace element analysis in plant material and ICP applications at Cornell University, Ithaca, NY-USA

21.09-11.10.2003     Short-term laboratory study at University of Western Australia, Perth-Australia

01.03-31.03.2004     Short-term laboratory study at Hohenheim University, Institute for Plant Nutrition, Stuttgart-Germany

08.09-19.09.2006     NATO Advanced Study Institute “Novel biotechnologies for biocontrol agent enhancement and management”, Perugia-Italy




BIO 335 Analytical Techniques

BIO 330 Environmental Plant Biology

BIO 543 Plant Stress Physiology







2002-2006 TÜBİTAK TOGTAG-2916: Tolerance to Low Temperature Stress and the Role of Antioxidative Defense Mechanisms in Tomato

2008-2011 TÜBİTAK 108T436: Physiological characterization of mechanisms involved in transport of zinc in to wheat grain

2013-2016 TÜBİTAK 113Z129: Effects of Magnesium and Potassium Nutrition on Transport of Photoassimilates under Elevated Carbon Dioxide Conditions







1993-1997 NATO-SFS/TU-Genotypes: Selection and Characterization of Cereal Genotypes with High Zinc Efficiency and Evaluation of Bioavailability of Zinc in Wheat for the Central Anatolia Region

1994-1997 GIARA-D223 (German Israel Agricultural Research Agreement): Role of Oxygen Radicals and Lipid Peroxidation in Drought Resistance in Wheat Genotypes

1996-1997 DANIDA/IFPRI: Evaluation of Micro Element Concentrations of Turkish and Other Origin Wheat Genotypes.

1996-1998 NATO-CP: Evaluation of Wheat Genotypes Resistant to Zn Deficiency and Localization of Genes Responsible from Resistance to Zinc Deficiency

2005-2007 HarvestPlus Biofortification Challenge Program: Improving Zinc and Protein Content of Wheat Grain by Introgression of Genes from its Wild Progenitor

2005-2009 EU-Project QUALIFO (subcontractor): Organic and Low Input Production Methods

2005-2008 CIMMYT-HarvestPlus: Agronomy-Related Approaches Improving Wheat Zinc Nutrition

2008-2010 HarvestPlus: Use of Zinc-Containing Fertilizers for Enriching Cereal Grains with Zinc and Improving Yield in Different Countries (1st Phase)

2011-2014 HarvestPlus: Use of Zinc-Containing Fertilizers for Enriching Cereal Grains with Zinc and Improving Yield in Different Countries (2nd Phase)

2010-2016 HarvestPlus: Improving Zinc and Protein Content of Wheat Grain by Introgression of Genes from its Wild Progenitor

2009-2014 European Comission: Improving Nutrient Efficiency in Major European Food, Feed and Biofuel Crops to Reduce the Negative Environmental Impact of Crop Production (Project NUE-crops)

2013-2018 European Comission: An integrated approach to diversify the genetic base, improve stress resistance, agronomic management and nutritional/processing quality of minor cereal crops for human nutrition in Europe.




1998-2001 TÜBİTAK (TOGTAG/TARP-2028): Screening of Phosphorus-Efficient Wheat Genotypes and Characterization of Morphological and Physiological Mechanisms Involved in Phosphorus Efficiency

2005-2006 Istanbul Municipality ISTAC Co.: Utilization of Municipal Solid Waste Compost as an Organic Soil Amendment

2005-2010 State Planning Organization: Exploitation of Wild Wheat to Improve Grain Concentration of Iron and Zinc in Modern Wheat

2006-2008 TÜBİTAK (105O637): Determination of Selenium Status of Soils and Cereals in Selected Regions in Turkey and Response of Wheat to Selenium Fertilizers

2006-2009 TÜBİTAK Kamu (105G148): Investigation of the IBB-Kemerburgaz Composting Facility Compost Product for its Utilization in Plant Production as an Organic Fertilizer

PUBLICATONS (Articles Published in International Refereed Journals and Indexed in SCI)


1.     Cakmak I, Ozturk L, Marschner H, Karanlık S, Ekiz H. 1996. Zinc-efficient wild grasses enhance release of phytosiderophores under zinc deficiency. J Plant Nutr 19, 551-563

2.     Cakmak I, Ozturk L, Eker S, Torun B, Kalfa HI, Yılmaz A. 1997. Concentration of zinc and activity of copper/zinc-superoxide dismutase in leaves of rye and wheat cultivars differing in sensitivity to zinc deficiency. J Plant Physiol 151, 91-95

3.     Cakmak I, Torun B, Erenoglu B, Ozturk L, Marschner H, Kalaycı M, Ekiz H, Yılmaz A. 1998. Morphological and physiological differences in the response of cereals to zinc deficiency. Euphytica 100, 349-357

4.     Kalaycı M, Torun B, Eker S, Aydın M, Ozturk L, Cakmak I. 1999. Grain yield, zinc efficiency and zinc concentration of wheat cultivars grown in a zinc-deficient calcareous soil in field and greenhouse. Field Crops Res 63, 87-98

5.     Cakmak I, Tolay I, Ozdemir A, Ozturk L, Kling CI .1999. Differences in zinc efficiency among and within diploid, tetraploid and hexaploid wheats. Ann Botany 84, 163-171

6.     Cakmak I, Welch RM, Hardt J, Norvell WA, Ozturk L, Kochian LV. 1999. Uptake and re-translocation of leaf-applied cadmium (109-Cd) in diploid, tetraploid and hexaploid wheats. J. Exp. Botany 51, 221-226

7.     Cakmak O, Ozturk L, Torun B, Ozkan H, Cakmak I. 2001. Tolerance of 65 durum wheat genotypes to zinc deficiency in a calcareous soil. J Plant Nutr 24, 1831-1847

8.     Torun B, Kalayci M, Ozturk L, Torun A, Aydin M, Cakmak I. 2003. Differences in shoot boron concentrations, leaf symptoms, and yield of Turkish barley cultivars grown on boron-toxic soil in field. J Plant Nutr 26, 1735-1747

9.     Dasgan HY, Ozturk L, Abak K, Cakmak I. 2003. Activities of iron-containing enzymes in leaves of two tomato genotypes differing in their resistance to Fe chlorosis. J Plant Nutr 26, 1997-2007

10. Ozturk L, Karanlik S, Ozkutlu F, Cakmak I, Kochian LV. 2003. Shoot biomass and zinc/cadmium uptake for hyperaccumulator and non-accumulator Thlaspi species in response to growth on a zinc-deficient calcareous soil. Plant Sci 164, 1095-1101

11. Hacisalihoglu G, Ozturk L, Cakmak I, Welch RM, Kochian LV. 2004. Genotypic Variation in Common Bean in Response to Zinc Deficiency in Calcarous Soil. Plant Soil 259, 71-83

12. Ozturk L, Eker S, Torun B, Cakmak I. 2005. Variation in phosphorus efficiency among 73 bread and durum wheat genotypes grown in a phosphorus-deficient calcareous soil. Plant Soil 269, 69-80

13. Ozturk L, Yazici MA, Yucel C, Torun A, Cekic C, Bagci A, Ozkan H, Braun HJ, Sayers Z, Cakmak I. 2006. Concentration and localization of zinc during seed development and germination in wheat. Physiol Plant 128, 144–152

14. Eker S, Ozturk L, Yazici A, Erenoglu B, Romheld V, Cakmak I. 2006. Foliar-applied glyphosate substantially reduced uptake and transport of iron and manganese in sunflower (Helianthus annuus L.) plants. J Agr Food Chem 54: 10019-10025

15. Ozkutlu F, Ozturk L, Erdem H, McLaughlin M, Cakmak I. 2007. Leaf-applied sodium chloride promotes cadmium accumulation in durum wheat grain. Plant Soil 290: 323-331

16. Distelfeld A, Cakmak I, Peleg Z, Ozturk L, Yazici AM, Budak H, Saranga Y, Fahima T. 2007. Multiple QTL-effects of wheat Gpc-B1 locus on grain protein and micronutrient concentrations. Physiol. Plant. 129: 635-643

17. Morgounov A, Gomez-Becerra HF, Abugalieva A, Dzhunusova M, Yessimbekova M, Muminjanov H, Zelenskiy Y, Ozturk L, Cakmak I. 2007. Iron and zinc grain density in common wheat grown in Central Asia. Euphytica 155: 193-203

18. Ozturk L, Yazici A, Eker S, Gokmen O, Rolmheld V, Cakmak I. 2008. Glyphosate inhibition of ferric reductase activity in iron deficient sunflower roots. New Phytol 177: 899-906

19. Peleg Z, Saranga Y, Yazici A, Fahima T, Ozturk L, Cakmak I. 2008. Grain zinc, iron and protein concentrations and zinc-efficiency in wild emmer wheat under contrasting irrigation regimes. Plant Soil 306: 57-67

20. Cebeci O, Kokturk B, Ergen N, Ozturk L, Cakmak I, Budak H. 2008. Differential expression of wheat transcriptomes in response to varying cadmium concentrations. Biol Plant 52: 703-708

21. Peleg Z, Cakmak I, Ozturk L, Yazici A, Jun Y, Budak H, Korol AB, Fahima T, Saranga Y. 2009. Quantitative trait loci conferring grain mineral nutrient concentrations in durum wheat x wild emmer wheat RIL population. Theor Appl Genet 119: 353-369

22. Cakmak I, Yazici A, Tutus Y, Ozturk L. 2009. Glyphosate reduced seed and leaf concentrations of calcium, manganese, magnesium, and iron in non-glyphosate resistant soybean. Eur J Agron 31: 114-119

23. Su YS, Ozturk L, Cakmak I, Budak H. 2009. Turfgrass species response exposed to increasing rates of glyphosate application. Eur J Agron 31: 120-125

24. Gomez-Becerra HF, Yazici A, Ozturk L, Budak H, Peleg Z, Morgounov A, Fahima T, Saranga Y, Cakmak I. 2010. Genetic variation and environmental stability of grain mineral nutrient concentrations in Triticum dicoccoides under five environments. Euphytica 171: 39-52

25. Kutman UB, Yildiz B, Ozturk L, Cakmak I. 2010. Biofortification of Durum Wheat with Zinc Through Soil and Foliar Applications of Nitrogen. Cereal Chem 87: 1-9

26. Chatzav M, Peleg Z, Ozturk L, Yazici A, Fahima T, Cakmak I, Saranga Y. 2010. Genetic diversity for grain nutrients in wild emmer wheat: potential for wheat improvement. Ann Botany 105: 1211-1220

27. Cakmak I, Kalayci M, Kaya Y, Torun AA, Aydin N, Wang Y, Arisoy Z, Erdem H, Yazici A, Gokmen O, Ozturk L, Horst WJ. 2010. Biofortification and Localization of Zinc in Wheat Grain. J Agr Food Chem 58: 9092-9102

28. Gomez-Becerra HF, Erdem, H, Yazici A, Tutus Y, Torun B, Ozturk L, Cakmak, I. 2010. Grain concentrations of protein and mineral nutrients in a large collection of spelt wheat grown under different environments. J Cereal Sci 52: 342-349

29. Kabaha K, Taralp A, Cakmak I, Ozturk L. 2011.Accelerated Hydrolysis Method To Estimate the Amino Acid Content of Wheat (Triticum durum Desf.) Flour Using Microwave Irradiation. J Agr Food Chem 59: 2958-2965

30. Cooper J, Sanderson R, Cakmak I, Ozturk L, Shotton P, Carmichael A, Haghighi RS, Tetard-Jones C, Volakakis N, Eyre M, Leifert C. 2011. Effect of Organic and Conventional Crop Rotation, Fertilization, and Crop Protection Practices on Metal Contents in Wheat (Triticum aestivum). J Agr Food Chem 59: 4715-4724

31. Aciksoz SB, Ozturk L, Gokmen OO, Romheld V, Cakmak I. 2011.Effect of nitrogen on root release of phytosiderophores and root uptake of Fe(III)-phytosiderophore in Fe-deficient wheat plants. Physiol Plant 142: 287-296

32. Durmaz E, Coruh C, Dinler G, Grusak MA, Peleg Z, Saranga Y, Fahima T, Yazici A, Ozturk L,Cakmak I, Budak H. 2011. Expression and Cellular Localization of ZIP1 Transporter Under Zinc Deficiency in Wild Emmer Wheat. Plant Mol Biol Rep 29: 582-596

33. Aciksoz SB, Yazici A, Ozturk L, Cakmak I. 2011.Biofortification of wheat with iron through soil and foliar application of nitrogen and iron fertilizers. Plant Soil 349: 215-225

34. Srednicka-Tober D, Baranski M, Gromadzka-Ostrowska J, Skwarlo-Sonta K, Rembialkowska E, Hajslova J, Schulzova V, Cakmak I, Ozturk L et al. 2013 Effect of Crop Protection and Fertilization Regimes Used in Organic and Conventional Production Systems on Feed Composition and Physiological Parameters in Rats. J Agr Food Chem 61: 1017-1029

35. Palmer MW, Cooper J, Tetard-Jones C, Srednicka-Tober D, Baranski M, Eyre M, Shotton PN, Volakakis N, Cakmak I, Ozturk L et al. 2013. The influence of organic and conventional fertilisation and crop protection practices, preceding crop, harvest year and weather conditions on yield and quality of potato (Solanum tuberosum) in a long-term management trial Eur J Agr 49: 83-92

36. Bilsborrow P, Cooper J, Tetard-Jones C, Srednicka-Tober D, Baraniski M, Eyre M, Schmidt C, Shotton P, Volakakis N, Cakmak I, Ozturk L et al. 2013. The effect of organic and conventional management on the yield and quality of wheat grown in a long-term field trial. Eur J Agr 51: 71-80

37. Aciksoz SB, Ozturk L, Yazici A, Cakmak I. 2014. Inclusion of urea in a (59)FeEDTA solution stimulated leaf penetration and translocation of Fe-59 within wheat plants. Physiol Plant 151: 348-357

38. Ova EA, Kutman UB, Ozturk L, Cakmak I. 2015. High phosphorus supply reduced zinc concentration of wheat in native soil but not in autoclaved soil or nutrient solution. Plant Soil 393: 147-162



(Updated September 24th 2015)